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Wales have suffered a double late blow with Liam Williams and Dafydd Jenkins dominated out of the Six Nations match in opposition to Italy on Saturday.
Blair Murray and Freddie Thomas will now begin the important thing fixture at Stadio Olimpico.
It is a serious blow for Warren Gatland to lose his most skilled again and one in every of his finest performing forwards for such an vital sport.
Veteran full-back Williams had been battling a knee damage sustained in opposition to France within the opening sport and missed coaching on Friday.
Scarlets wing Murray – who performed all three Tests throughout the autumn, together with at full-back in opposition to South Africa – wins solely his fifth cap.
Jenkins, who made the second most tackles of any participant within the Six Nations final week, has been dominated out by means of sickness.
Gloucester lock Thomas makes his first Wales begin having made his debut in opposition to South Africa in November.
His Kingsholm club-mate Josh Hathaway and Cardiff lock Teddy Williams have been promoted to the bench.
Wales misplaced 43-0 in opposition to France in Paris of their opening Six Nations sport, heightening the issues for Gatland’s squad after 13 consecutive Test defeats.
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